##gff-version 2 (GTF)  piRNAome Rattus norvegicus piRNA clusters (known + de novo); 1-based
chr1	piRNAome	piRNA_cluster	55467003	55485042	.	+	.	cluster_id "piRO-rno-C00000088"; cluster_type "known"; n_piRNA "64";
chr1	piRNAome	piRNA_cluster	134408321	134431776	.	-	.	cluster_id "piRO-rno-C00000089"; cluster_type "known"; n_piRNA "120";
chr1	piRNAome	piRNA_cluster	135136992	135227807	.	.	.	cluster_id "piRO-rno-C00000090"; cluster_type "known"; n_piRNA "2673";
chr1	piRNAome	piRNA_cluster	196691918	196702337	.	-	.	cluster_id "piRO-rno-C00000091"; cluster_type "known"; n_piRNA "190";
chr1	piRNAome	piRNA_cluster	221201343	221204144	.	-	.	cluster_id "piRO-rno-C00000092"; cluster_type "known"; n_piRNA "17";
chr1	piRNAome	piRNA_cluster	255891256	255901398	.	-	.	cluster_id "piRO-rno-C00000093"; cluster_type "known"; n_piRNA "225";
chr2	piRNAome	piRNA_cluster	44870878	44890159	.	-	.	cluster_id "piRO-rno-C00000094"; cluster_type "known"; n_piRNA "34";
chr2	piRNAome	piRNA_cluster	105273767	105285665	.	-	.	cluster_id "piRO-rno-C00000095"; cluster_type "known"; n_piRNA "61";
chr2	piRNAome	piRNA_cluster	228642007	228665289	.	-	.	cluster_id "piRO-rno-C00000096"; cluster_type "known"; n_piRNA "153";
chr3	piRNAome	piRNA_cluster	21819338	21822743	.	-	.	cluster_id "piRO-rno-C00000097"; cluster_type "known"; n_piRNA "8";
chr3	piRNAome	piRNA_cluster	81432310	81505102	.	.	.	cluster_id "piRO-rno-C00000098"; cluster_type "known"; n_piRNA "2819";
chr3	piRNAome	piRNA_cluster	82069488	82081047	.	-	.	cluster_id "piRO-rno-C00000099"; cluster_type "known"; n_piRNA "8";
chr3	piRNAome	piRNA_cluster	145906398	145916788	.	-	.	cluster_id "piRO-rno-C00000100"; cluster_type "known"; n_piRNA "38";
chr3	piRNAome	piRNA_cluster	146179610	146203392	.	-	.	cluster_id "piRO-rno-C00000101"; cluster_type "known"; n_piRNA "348";
chr3	piRNAome	piRNA_cluster	146854160	146862169	.	-	.	cluster_id "piRO-rno-C00000102"; cluster_type "known"; n_piRNA "187";
chr4	piRNAome	piRNA_cluster	112635817	112647791	.	-	.	cluster_id "piRO-rno-C00000103"; cluster_type "known"; n_piRNA "275";
chr4	piRNAome	piRNA_cluster	116827665	116838964	.	-	.	cluster_id "piRO-rno-C00000104"; cluster_type "known"; n_piRNA "490";
chr4	piRNAome	piRNA_cluster	116879729	116891640	.	-	.	cluster_id "piRO-rno-C00000105"; cluster_type "known"; n_piRNA "106";
chr4	piRNAome	piRNA_cluster	119868390	119902751	.	-	.	cluster_id "piRO-rno-C00000106"; cluster_type "known"; n_piRNA "583";
chr4	piRNAome	piRNA_cluster	160341300	160417256	.	.	.	cluster_id "piRO-rno-C00000107"; cluster_type "known"; n_piRNA "2968";
chr5	piRNAome	piRNA_cluster	24771284	24801835	.	-	.	cluster_id "piRO-rno-C00000108"; cluster_type "known"; n_piRNA "280";
chr5	piRNAome	piRNA_cluster	62129909	62141245	.	-	.	cluster_id "piRO-rno-C00000109"; cluster_type "known"; n_piRNA "38";
chr5	piRNAome	piRNA_cluster	74432491	74443282	.	-	.	cluster_id "piRO-rno-C00000110"; cluster_type "known"; n_piRNA "272";
chr5	piRNAome	piRNA_cluster	77988856	77990457	.	-	.	cluster_id "piRO-rno-C00000111"; cluster_type "known"; n_piRNA "47";
chr5	piRNAome	piRNA_cluster	78071437	78071572	.	-	.	cluster_id "piRO-rno-C00000112"; cluster_type "known"; n_piRNA "9";
chr5	piRNAome	piRNA_cluster	113209917	113260416	.	-	.	cluster_id "piRO-rno-C00000113"; cluster_type "known"; n_piRNA "1535";
chr1	piRNAome	piRNA_cluster	224998540	225000025	.	+	.	cluster_id "piRO-rno-C00000114"; cluster_type "known"; n_piRNA "10";
chr5	piRNAome	piRNA_cluster	141510813	141517990	.	-	.	cluster_id "piRO-rno-C00000115"; cluster_type "known"; n_piRNA "95";
chr5	piRNAome	piRNA_cluster	141546917	141551039	.	-	.	cluster_id "piRO-rno-C00000116"; cluster_type "known"; n_piRNA "7";
chr6	piRNAome	piRNA_cluster	94180797	94201896	.	.	.	cluster_id "piRO-rno-C00000117"; cluster_type "known"; n_piRNA "71";
chr6	piRNAome	piRNA_cluster	122356429	122438743	.	.	.	cluster_id "piRO-rno-C00000118"; cluster_type "known"; n_piRNA "2513";
chr7	piRNAome	piRNA_cluster	18512266	18518869	.	-	.	cluster_id "piRO-rno-C00000119"; cluster_type "known"; n_piRNA "7";
chr7	piRNAome	piRNA_cluster	97844634	97872443	.	-	.	cluster_id "piRO-rno-C00000120"; cluster_type "known"; n_piRNA "118";
chr7	piRNAome	piRNA_cluster	99535012	99594802	.	.	.	cluster_id "piRO-rno-C00000121"; cluster_type "known"; n_piRNA "1413";
chr7	piRNAome	piRNA_cluster	119838428	119892638	.	-	.	cluster_id "piRO-rno-C00000122"; cluster_type "known"; n_piRNA "602";
chr2	piRNAome	piRNA_cluster	121390075	121395810	.	+	.	cluster_id "piRO-rno-C00000123"; cluster_type "known"; n_piRNA "41";
chr7	piRNAome	piRNA_cluster	124201914	124214567	.	-	.	cluster_id "piRO-rno-C00000124"; cluster_type "known"; n_piRNA "146";
chr7	piRNAome	piRNA_cluster	144813315	144826223	.	-	.	cluster_id "piRO-rno-C00000125"; cluster_type "known"; n_piRNA "17";
chr8	piRNAome	piRNA_cluster	39755527	39760632	.	-	.	cluster_id "piRO-rno-C00000126"; cluster_type "known"; n_piRNA "30";
chr8	piRNAome	piRNA_cluster	48482475	48486934	.	-	.	cluster_id "piRO-rno-C00000127"; cluster_type "known"; n_piRNA "48";
chr8	piRNAome	piRNA_cluster	58784252	58836787	.	.	.	cluster_id "piRO-rno-C00000128"; cluster_type "known"; n_piRNA "2017";
chr8	piRNAome	piRNA_cluster	73534002	73591520	.	.	.	cluster_id "piRO-rno-C00000129"; cluster_type "known"; n_piRNA "2061";
chr9	piRNAome	piRNA_cluster	12693087	12695784	.	-	.	cluster_id "piRO-rno-C00000130"; cluster_type "known"; n_piRNA "18";
chr9	piRNAome	piRNA_cluster	98540441	98544072	.	-	.	cluster_id "piRO-rno-C00000131"; cluster_type "known"; n_piRNA "71";
chr9	piRNAome	piRNA_cluster	100148580	100152511	.	-	.	cluster_id "piRO-rno-C00000132"; cluster_type "known"; n_piRNA "140";
chr9	piRNAome	piRNA_cluster	114166797	114190725	.	.	.	cluster_id "piRO-rno-C00000133"; cluster_type "known"; n_piRNA "138";
chr9	piRNAome	piRNA_cluster	114337242	114341138	.	-	.	cluster_id "piRO-rno-C00000134"; cluster_type "known"; n_piRNA "8";
chr10	piRNAome	piRNA_cluster	36441747	36444185	.	-	.	cluster_id "piRO-rno-C00000135"; cluster_type "known"; n_piRNA "63";
chr10	piRNAome	piRNA_cluster	96659537	96662296	.	-	.	cluster_id "piRO-rno-C00000136"; cluster_type "known"; n_piRNA "9";
chr11	piRNAome	piRNA_cluster	84231571	84240821	.	-	.	cluster_id "piRO-rno-C00000137"; cluster_type "known"; n_piRNA "20";
chr11	piRNAome	piRNA_cluster	87085795	87133726	.	.	.	cluster_id "piRO-rno-C00000138"; cluster_type "known"; n_piRNA "441";
chr12	piRNAome	piRNA_cluster	6503565	6543796	.	-	.	cluster_id "piRO-rno-C00000139"; cluster_type "known"; n_piRNA "1770";
chr12	piRNAome	piRNA_cluster	12830314	12837663	.	-	.	cluster_id "piRO-rno-C00000140"; cluster_type "known"; n_piRNA "31";
chr12	piRNAome	piRNA_cluster	22778567	22796229	.	-	.	cluster_id "piRO-rno-C00000141"; cluster_type "known"; n_piRNA "186";
chr12	piRNAome	piRNA_cluster	30022223	30033734	.	-	.	cluster_id "piRO-rno-C00000142"; cluster_type "known"; n_piRNA "92";
chr12	piRNAome	piRNA_cluster	49058818	49092468	.	.	.	cluster_id "piRO-rno-C00000143"; cluster_type "known"; n_piRNA "961";
chr13	piRNAome	piRNA_cluster	76021745	76034791	.	-	.	cluster_id "piRO-rno-C00000144"; cluster_type "known"; n_piRNA "20";
chr14	piRNAome	piRNA_cluster	3726469	3728001	.	-	.	cluster_id "piRO-rno-C00000145"; cluster_type "known"; n_piRNA "9";
chr14	piRNAome	piRNA_cluster	41061885	41073545	.	-	.	cluster_id "piRO-rno-C00000146"; cluster_type "known"; n_piRNA "125";
chr3	piRNAome	piRNA_cluster	120255760	120272530	.	+	.	cluster_id "piRO-rno-C00000147"; cluster_type "known"; n_piRNA "521";
chr14	piRNAome	piRNA_cluster	85763743	85772751	.	-	.	cluster_id "piRO-rno-C00000148"; cluster_type "known"; n_piRNA "24";
chr15	piRNAome	piRNA_cluster	199695	279657	.	-	.	cluster_id "piRO-rno-C00000149"; cluster_type "known"; n_piRNA "946";
chr16	piRNAome	piRNA_cluster	48474635	48479056	.	-	.	cluster_id "piRO-rno-C00000150"; cluster_type "known"; n_piRNA "9";
chr17	piRNAome	piRNA_cluster	11756591	11761269	.	-	.	cluster_id "piRO-rno-C00000151"; cluster_type "known"; n_piRNA "37";
chr17	piRNAome	piRNA_cluster	11785768	11794290	.	-	.	cluster_id "piRO-rno-C00000152"; cluster_type "known"; n_piRNA "37";
chr17	piRNAome	piRNA_cluster	11818524	11838494	.	-	.	cluster_id "piRO-rno-C00000153"; cluster_type "known"; n_piRNA "262";
chr17	piRNAome	piRNA_cluster	13119618	13143115	.	-	.	cluster_id "piRO-rno-C00000154"; cluster_type "known"; n_piRNA "111";
chr3	piRNAome	piRNA_cluster	145487801	145497042	.	+	.	cluster_id "piRO-rno-C00000155"; cluster_type "known"; n_piRNA "138";
chr17	piRNAome	piRNA_cluster	15257250	15271754	.	-	.	cluster_id "piRO-rno-C00000156"; cluster_type "known"; n_piRNA "37";
chr17	piRNAome	piRNA_cluster	42159986	42161056	.	-	.	cluster_id "piRO-rno-C00000157"; cluster_type "known"; n_piRNA "17";
chr18	piRNAome	piRNA_cluster	25369584	25371628	.	-	.	cluster_id "piRO-rno-C00000158"; cluster_type "known"; n_piRNA "11";
chr18	piRNAome	piRNA_cluster	62772963	62773002	.	-	.	cluster_id "piRO-rno-C00000159"; cluster_type "known"; n_piRNA "11";
chr19	piRNAome	piRNA_cluster	41520839	41534388	.	-	.	cluster_id "piRO-rno-C00000160"; cluster_type "known"; n_piRNA "638";
chr20	piRNAome	piRNA_cluster	6526189	6527102	.	-	.	cluster_id "piRO-rno-C00000161"; cluster_type "known"; n_piRNA "103";
chr20	piRNAome	piRNA_cluster	6599550	6698425	.	.	.	cluster_id "piRO-rno-C00000162"; cluster_type "known"; n_piRNA "5781";
chr20	piRNAome	piRNA_cluster	13726210	13759541	.	-	.	cluster_id "piRO-rno-C00000163"; cluster_type "known"; n_piRNA "1317";
chr3	piRNAome	piRNA_cluster	145596470	145614248	.	+	.	cluster_id "piRO-rno-C00000164"; cluster_type "known"; n_piRNA "166";
chr20	piRNAome	piRNA_cluster	18459762	18467828	.	-	.	cluster_id "piRO-rno-C00000165"; cluster_type "known"; n_piRNA "14";
chrX	piRNAome	piRNA_cluster	151252838	151287819	.	-	.	cluster_id "piRO-rno-C00000166"; cluster_type "known"; n_piRNA "24";
chr1	piRNAome	piRNA_cluster	55707618	55724527	.	+	.	cluster_id "piRO-rno-C00000167"; cluster_type "known"; n_piRNA "146";
chr3	piRNAome	piRNA_cluster	146894816	146895892	.	+	.	cluster_id "piRO-rno-C00000168"; cluster_type "known"; n_piRNA "6";
chr3	piRNAome	piRNA_cluster	146929112	146945819	.	+	.	cluster_id "piRO-rno-C00000169"; cluster_type "known"; n_piRNA "583";
chr4	piRNAome	piRNA_cluster	153438123	153441125	.	+	.	cluster_id "piRO-rno-C00000170"; cluster_type "known"; n_piRNA "31";
chr4	piRNAome	piRNA_cluster	160838806	160840485	.	+	.	cluster_id "piRO-rno-C00000171"; cluster_type "known"; n_piRNA "56";
chr4	piRNAome	piRNA_cluster	160965937	160967250	.	+	.	cluster_id "piRO-rno-C00000172"; cluster_type "known"; n_piRNA "17";
chr4	piRNAome	piRNA_cluster	160993882	160996868	.	+	.	cluster_id "piRO-rno-C00000173"; cluster_type "known"; n_piRNA "33";
chr4	piRNAome	piRNA_cluster	161757663	161776001	.	+	.	cluster_id "piRO-rno-C00000174"; cluster_type "known"; n_piRNA "143";
chr5	piRNAome	piRNA_cluster	24975749	24991176	.	+	.	cluster_id "piRO-rno-C00000175"; cluster_type "known"; n_piRNA "16";
chr1	piRNAome	piRNA_cluster	56001058	56005142	.	+	.	cluster_id "piRO-rno-C00000176"; cluster_type "known"; n_piRNA "48";
chr5	piRNAome	piRNA_cluster	78073596	78082458	.	+	.	cluster_id "piRO-rno-C00000177"; cluster_type "known"; n_piRNA "821";
chr5	piRNAome	piRNA_cluster	153976546	153979809	.	+	.	cluster_id "piRO-rno-C00000178"; cluster_type "known"; n_piRNA "302";
chr6	piRNAome	piRNA_cluster	27916733	27918620	.	+	.	cluster_id "piRO-rno-C00000179"; cluster_type "known"; n_piRNA "7";
chr7	piRNAome	piRNA_cluster	20226164	20251144	.	+	.	cluster_id "piRO-rno-C00000180"; cluster_type "known"; n_piRNA "393";
chr7	piRNAome	piRNA_cluster	20394103	20396042	.	+	.	cluster_id "piRO-rno-C00000181"; cluster_type "known"; n_piRNA "19";
chr7	piRNAome	piRNA_cluster	22995875	23010813	.	+	.	cluster_id "piRO-rno-C00000182"; cluster_type "known"; n_piRNA "16";
chr1	piRNAome	piRNA_cluster	108279071	108283886	.	+	.	cluster_id "piRO-rno-C00000183"; cluster_type "known"; n_piRNA "22";
chr7	piRNAome	piRNA_cluster	115874499	115895444	.	+	.	cluster_id "piRO-rno-C00000184"; cluster_type "known"; n_piRNA "946";
chr8	piRNAome	piRNA_cluster	118803372	118815737	.	+	.	cluster_id "piRO-rno-C00000185"; cluster_type "known"; n_piRNA "9";
chr9	piRNAome	piRNA_cluster	13830560	13831662	.	+	.	cluster_id "piRO-rno-C00000186"; cluster_type "known"; n_piRNA "91";
chr9	piRNAome	piRNA_cluster	98672615	98689102	.	+	.	cluster_id "piRO-rno-C00000187"; cluster_type "known"; n_piRNA "80";
chr9	piRNAome	piRNA_cluster	113969452	114014246	.	.	.	cluster_id "piRO-rno-C00000188"; cluster_type "known"; n_piRNA "886";
chr9	piRNAome	piRNA_cluster	114341830	114368001	.	+	.	cluster_id "piRO-rno-C00000189"; cluster_type "known"; n_piRNA "98";
chr10	piRNAome	piRNA_cluster	15724755	15731561	.	+	.	cluster_id "piRO-rno-C00000190"; cluster_type "known"; n_piRNA "8";
chr10	piRNAome	piRNA_cluster	83572558	83577729	.	+	.	cluster_id "piRO-rno-C00000191"; cluster_type "known"; n_piRNA "25";
chr10	piRNAome	piRNA_cluster	89517275	89528192	.	+	.	cluster_id "piRO-rno-C00000192"; cluster_type "known"; n_piRNA "34";
chr10	piRNAome	piRNA_cluster	91514244	91543643	.	+	.	cluster_id "piRO-rno-C00000193"; cluster_type "known"; n_piRNA "654";
chr12	piRNAome	piRNA_cluster	47515606	47551411	.	+	.	cluster_id "piRO-rno-C00000194"; cluster_type "known"; n_piRNA "614";
chr12	piRNAome	piRNA_cluster	49307282	49325935	.	+	.	cluster_id "piRO-rno-C00000195"; cluster_type "known"; n_piRNA "277";
chr13	piRNAome	piRNA_cluster	41504515	41507360	.	+	.	cluster_id "piRO-rno-C00000196"; cluster_type "known"; n_piRNA "128";
chr13	piRNAome	piRNA_cluster	76035744	76052912	.	+	.	cluster_id "piRO-rno-C00000197"; cluster_type "known"; n_piRNA "125";
chr15	piRNAome	piRNA_cluster	84765936	84768424	.	+	.	cluster_id "piRO-rno-C00000198"; cluster_type "known"; n_piRNA "67";
chr17	piRNAome	piRNA_cluster	14335254	14336932	.	+	.	cluster_id "piRO-rno-C00000199"; cluster_type "known"; n_piRNA "8";
chr17	piRNAome	piRNA_cluster	14401463	14404617	.	+	.	cluster_id "piRO-rno-C00000200"; cluster_type "known"; n_piRNA "66";
chr1	piRNAome	piRNA_cluster	134431931	134474329	.	+	.	cluster_id "piRO-rno-C00000201"; cluster_type "known"; n_piRNA "1544";
chr17	piRNAome	piRNA_cluster	14983128	15006048	.	+	.	cluster_id "piRO-rno-C00000202"; cluster_type "known"; n_piRNA "61";
chr17	piRNAome	piRNA_cluster	15210325	15256486	.	+	.	cluster_id "piRO-rno-C00000203"; cluster_type "known"; n_piRNA "388";
chr19	piRNAome	piRNA_cluster	16236248	16253314	.	+	.	cluster_id "piRO-rno-C00000204"; cluster_type "known"; n_piRNA "588";
chr19	piRNAome	piRNA_cluster	41935384	41937674	.	+	.	cluster_id "piRO-rno-C00000205"; cluster_type "known"; n_piRNA "7";
chr19	piRNAome	piRNA_cluster	49744968	49750598	.	+	.	cluster_id "piRO-rno-C00000206"; cluster_type "known"; n_piRNA "7";
chr20	piRNAome	piRNA_cluster	23333081	23336167	.	+	.	cluster_id "piRO-rno-C00000207"; cluster_type "known"; n_piRNA "43";
chr20	piRNAome	piRNA_cluster	28646772	28657960	.	+	.	cluster_id "piRO-rno-C00000208"; cluster_type "known"; n_piRNA "126";
chr20	piRNAome	piRNA_cluster	32269812	32316115	.	+	.	cluster_id "piRO-rno-C00000209"; cluster_type "known"; n_piRNA "573";
chr20	piRNAome	piRNA_cluster	33872384	33873608	.	+	.	cluster_id "piRO-rno-C00000210"; cluster_type "known"; n_piRNA "9";
chrX	piRNAome	piRNA_cluster	73080950	73088770	.	+	.	cluster_id "piRO-rno-C00000211"; cluster_type "known"; n_piRNA "37";
chrX	piRNAome	piRNA_cluster	100582066	100587255	.	+	.	cluster_id "piRO-rno-C00000212"; cluster_type "known"; n_piRNA "189";
chrX	piRNAome	piRNA_cluster	107054195	107054260	.	+	.	cluster_id "piRO-rno-C00000213"; cluster_type "known"; n_piRNA "16";
chrX	piRNAome	piRNA_cluster	155330302	155344262	.	+	.	cluster_id "piRO-rno-C00000214"; cluster_type "known"; n_piRNA "10";
chr1	piRNAome	piRNA_cluster	14132498	14148017	.	-	.	cluster_id "piRO-rno-C00000215"; cluster_type "known"; n_piRNA "224";
chr1	piRNAome	piRNA_cluster	14181674	14201257	.	-	.	cluster_id "piRO-rno-C00000216"; cluster_type "known"; n_piRNA "244";
chr1	piRNAome	piRNA_cluster	54183117	54212715	.	-	.	cluster_id "piRO-rno-C00000217"; cluster_type "known"; n_piRNA "577";
chr1	piRNAome	piRNA_cluster	54888793	54901174	.	-	.	cluster_id "piRO-rno-C00000218"; cluster_type "known"; n_piRNA "239";
chr1	piRNAome	piRNA_cluster	55197602	55205690	.	-	.	cluster_id "piRO-rno-C00000219"; cluster_type "known"; n_piRNA "41";
chr1	piRNAome	piRNA_cluster	97767401	97781557	.	-	.	cluster_id "piRO-rno-C00000220"; cluster_type "known"; n_piRNA "75";
chr1	piRNAome	piRNA_cluster	107557396	107563130	.	-	.	cluster_id "piRO-rno-C00000221"; cluster_type "known"; n_piRNA "179";
chr1	piRNAome	piRNA_cluster	108830213	108840576	.	-	.	cluster_id "piRO-rno-C00000222"; cluster_type "known"; n_piRNA "19";
chr1	piRNAome	piRNA_cluster	109517534	109522453	.	-	.	cluster_id "piRO-rno-C00000223"; cluster_type "known"; n_piRNA "150";
chr1	piRNAome	piRNA_cluster	130981958	131050386	.	.	.	cluster_id "piRO-rno-C00000224"; cluster_type "known"; n_piRNA "1398";
chr1	piRNAome	piRNA_cluster	11903744	11909536	.	-	.	cluster_id "piRO-rno-C00000001"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "97"; poisson_pvalue "0.00e+00";
chr1	piRNAome	piRNA_cluster	14148564	14162642	.	-	.	cluster_id "piRO-rno-C00000002"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "10"; poisson_pvalue "6.54e-11";
chr1	piRNAome	piRNA_cluster	15798030	15799044	.	-	.	cluster_id "piRO-rno-C00000003"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "1.62e-12";
chr1	piRNAome	piRNA_cluster	54181112	54187893	.	-	.	cluster_id "piRO-rno-C00000004"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "9"; poisson_pvalue "2.50e-12";
chr1	piRNAome	piRNA_cluster	54209139	54214012	.	-	.	cluster_id "piRO-rno-C00000005"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "9"; poisson_pvalue "1.35e-13";
chr1	piRNAome	piRNA_cluster	54885196	54888699	.	-	.	cluster_id "piRO-rno-C00000006"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "2.56e-09";
chr1	piRNAome	piRNA_cluster	54931908	54935058	.	-	.	cluster_id "piRO-rno-C00000007"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "11"; poisson_pvalue "0.00e+00";
chr1	piRNAome	piRNA_cluster	86781109	86793672	.	-	.	cluster_id "piRO-rno-C00000008"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "7"; poisson_pvalue "2.43e-07";
chr1	piRNAome	piRNA_cluster	99054646	99056520	.	-	.	cluster_id "piRO-rno-C00000009"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "6"; poisson_pvalue "6.29e-11";
chr1	piRNAome	piRNA_cluster	107559314	107563250	.	-	.	cluster_id "piRO-rno-C00000010"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "5.10e-09";
chr1	piRNAome	piRNA_cluster	107580343	107583122	.	-	.	cluster_id "piRO-rno-C00000011"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "5"; poisson_pvalue "4.39e-08";
chr1	piRNAome	piRNA_cluster	108825061	108830281	.	-	.	cluster_id "piRO-rno-C00000012"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "26"; poisson_pvalue "0.00e+00";
chr1	piRNAome	piRNA_cluster	109515912	109521606	.	-	.	cluster_id "piRO-rno-C00000013"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "4.45e-08";
chr1	piRNAome	piRNA_cluster	11968863	11974363	.	+	.	cluster_id "piRO-rno-C00000014"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "84"; poisson_pvalue "0.00e+00";
chr1	piRNAome	piRNA_cluster	50808310	50808880	.	+	.	cluster_id "piRO-rno-C00000015"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "5"; poisson_pvalue "1.70e-11";
chr1	piRNAome	piRNA_cluster	55397479	55406113	.	+	.	cluster_id "piRO-rno-C00000016"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "21"; poisson_pvalue "0.00e+00";
chr1	piRNAome	piRNA_cluster	55447854	55453708	.	+	.	cluster_id "piRO-rno-C00000017"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "9"; poisson_pvalue "6.84e-13";
chr1	piRNAome	piRNA_cluster	55461524	55467958	.	+	.	cluster_id "piRO-rno-C00000018"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "9"; poisson_pvalue "1.58e-12";
chr1	piRNAome	piRNA_cluster	55493661	55499704	.	+	.	cluster_id "piRO-rno-C00000019"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "10"; poisson_pvalue "1.75e-14";
chr1	piRNAome	piRNA_cluster	56001920	56007771	.	+	.	cluster_id "piRO-rno-C00000020"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "5.21e-08";
chr1	piRNAome	piRNA_cluster	56023933	56025619	.	+	.	cluster_id "piRO-rno-C00000021"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "8"; poisson_pvalue "1.78e-15";
chr1	piRNAome	piRNA_cluster	108266087	108267993	.	+	.	cluster_id "piRO-rno-C00000022"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "6.96e-11";
chr1	piRNAome	piRNA_cluster	108275527	108278988	.	+	.	cluster_id "piRO-rno-C00000023"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "39"; poisson_pvalue "0.00e+00";
chr1	piRNAome	piRNA_cluster	109472402	109474912	.	+	.	cluster_id "piRO-rno-C00000024"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "28"; poisson_pvalue "0.00e+00";
chr1	piRNAome	piRNA_cluster	134474334	134478351	.	+	.	cluster_id "piRO-rno-C00000025"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "5.75e-09";
chr1	piRNAome	piRNA_cluster	222547357	222549768	.	+	.	cluster_id "piRO-rno-C00000026"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "5"; poisson_pvalue "2.18e-08";
chr10	piRNAome	piRNA_cluster	12852781	12854414	.	-	.	cluster_id "piRO-rno-C00000027"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "7"; poisson_pvalue "1.11e-16";
chr10	piRNAome	piRNA_cluster	55617326	55621356	.	+	.	cluster_id "piRO-rno-C00000028"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "9"; poisson_pvalue "0.00e+00";
chr10	piRNAome	piRNA_cluster	59644500	59647329	.	+	.	cluster_id "piRO-rno-C00000029"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "5"; poisson_pvalue "1.70e-10";
chr11	piRNAome	piRNA_cluster	84170325	84175217	.	-	.	cluster_id "piRO-rno-C00000030"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "36"; poisson_pvalue "0.00e+00";
chr11	piRNAome	piRNA_cluster	84243540	84247151	.	-	.	cluster_id "piRO-rno-C00000031"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "32"; poisson_pvalue "0.00e+00";
chr12	piRNAome	piRNA_cluster	30034563	30039531	.	+	.	cluster_id "piRO-rno-C00000032"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "5"; poisson_pvalue "4.66e-05";
chr13	piRNAome	piRNA_cluster	89439569	89440603	.	-	.	cluster_id "piRO-rno-C00000033"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "9"; poisson_pvalue "0.00e+00";
chr13	piRNAome	piRNA_cluster	91161469	91162518	.	-	.	cluster_id "piRO-rno-C00000034"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "8"; poisson_pvalue "0.00e+00";
chr14	piRNAome	piRNA_cluster	34017984	34022063	.	-	.	cluster_id "piRO-rno-C00000035"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "8"; poisson_pvalue "0.00e+00";
chr14	piRNAome	piRNA_cluster	85002283	85011278	.	-	.	cluster_id "piRO-rno-C00000036"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "5"; poisson_pvalue "1.49e-10";
chr14	piRNAome	piRNA_cluster	46517688	46526098	.	+	.	cluster_id "piRO-rno-C00000037"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "68"; poisson_pvalue "0.00e+00";
chr14	piRNAome	piRNA_cluster	46636768	46649165	.	+	.	cluster_id "piRO-rno-C00000038"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "11"; poisson_pvalue "0.00e+00";
chr15	piRNAome	piRNA_cluster	28015588	28017624	.	+	.	cluster_id "piRO-rno-C00000039"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "5"; poisson_pvalue "2.49e-11";
chr16	piRNAome	piRNA_cluster	81951475	81964911	.	-	.	cluster_id "piRO-rno-C00000040"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "5"; poisson_pvalue "3.88e-12";
chr16	piRNAome	piRNA_cluster	71677775	71680058	.	+	.	cluster_id "piRO-rno-C00000041"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "5"; poisson_pvalue "5.55e-16";
chr17	piRNAome	piRNA_cluster	11795494	11801762	.	-	.	cluster_id "piRO-rno-C00000042"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "9"; poisson_pvalue "5.55e-16";
chr17	piRNAome	piRNA_cluster	14432927	14434609	.	-	.	cluster_id "piRO-rno-C00000043"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "7"; poisson_pvalue "5.55e-16";
chr17	piRNAome	piRNA_cluster	14331200	14334612	.	+	.	cluster_id "piRO-rno-C00000044"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "9"; poisson_pvalue "0.00e+00";
chr17	piRNAome	piRNA_cluster	14397975	14405417	.	+	.	cluster_id "piRO-rno-C00000045"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "10"; poisson_pvalue "0.00e+00";
chr17	piRNAome	piRNA_cluster	15006186	15008174	.	+	.	cluster_id "piRO-rno-C00000046"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "6"; poisson_pvalue "4.69e-13";
chr17	piRNAome	piRNA_cluster	45635763	45637332	.	+	.	cluster_id "piRO-rno-C00000047"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "11"; poisson_pvalue "0.00e+00";
chr18	piRNAome	piRNA_cluster	60882873	60917695	.	-	.	cluster_id "piRO-rno-C00000048"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "642"; poisson_pvalue "0.00e+00";
chr18	piRNAome	piRNA_cluster	62758310	62793363	.	-	.	cluster_id "piRO-rno-C00000049"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "701"; poisson_pvalue "0.00e+00";
chr18	piRNAome	piRNA_cluster	70970450	70971300	.	+	.	cluster_id "piRO-rno-C00000050"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "7"; poisson_pvalue "0.00e+00";
chr2	piRNAome	piRNA_cluster	105285632	105288005	.	-	.	cluster_id "piRO-rno-C00000051"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "5"; poisson_pvalue "3.81e-13";
chr2	piRNAome	piRNA_cluster	105294768	105297055	.	-	.	cluster_id "piRO-rno-C00000052"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "5"; poisson_pvalue "3.17e-13";
chr2	piRNAome	piRNA_cluster	105332227	105349302	.	-	.	cluster_id "piRO-rno-C00000053"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "66"; poisson_pvalue "0.00e+00";
chr2	piRNAome	piRNA_cluster	105361580	105381076	.	-	.	cluster_id "piRO-rno-C00000054"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "109"; poisson_pvalue "0.00e+00";
chr2	piRNAome	piRNA_cluster	105387630	105395275	.	-	.	cluster_id "piRO-rno-C00000055"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "25"; poisson_pvalue "0.00e+00";
chr2	piRNAome	piRNA_cluster	105473942	105476280	.	-	.	cluster_id "piRO-rno-C00000056"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "9"; poisson_pvalue "0.00e+00";
chr2	piRNAome	piRNA_cluster	105513102	105538268	.	-	.	cluster_id "piRO-rno-C00000057"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "73"; poisson_pvalue "0.00e+00";
chr2	piRNAome	piRNA_cluster	105550820	105571289	.	-	.	cluster_id "piRO-rno-C00000058"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "85"; poisson_pvalue "0.00e+00";
chr2	piRNAome	piRNA_cluster	105576345	105584215	.	-	.	cluster_id "piRO-rno-C00000059"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "13"; poisson_pvalue "0.00e+00";
chr2	piRNAome	piRNA_cluster	105858744	105869331	.	-	.	cluster_id "piRO-rno-C00000060"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "18"; poisson_pvalue "0.00e+00";
chr2	piRNAome	piRNA_cluster	198630171	198638978	.	-	.	cluster_id "piRO-rno-C00000061"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "30"; poisson_pvalue "0.00e+00";
chr2	piRNAome	piRNA_cluster	124054735	124060660	.	+	.	cluster_id "piRO-rno-C00000062"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "6"; poisson_pvalue "1.30e-13";
chr3	piRNAome	piRNA_cluster	145916913	145922333	.	-	.	cluster_id "piRO-rno-C00000063"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "21"; poisson_pvalue "0.00e+00";
chr3	piRNAome	piRNA_cluster	146204255	146206438	.	-	.	cluster_id "piRO-rno-C00000064"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "9"; poisson_pvalue "0.00e+00";
chr3	piRNAome	piRNA_cluster	146849666	146854145	.	-	.	cluster_id "piRO-rno-C00000065"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "14"; poisson_pvalue "0.00e+00";
chr4	piRNAome	piRNA_cluster	117647014	117663099	.	-	.	cluster_id "piRO-rno-C00000066"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "101"; poisson_pvalue "0.00e+00";
chr4	piRNAome	piRNA_cluster	117829999	117838134	.	-	.	cluster_id "piRO-rno-C00000067"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "106"; poisson_pvalue "0.00e+00";
chr4	piRNAome	piRNA_cluster	161452367	161460100	.	-	.	cluster_id "piRO-rno-C00000068"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "13"; poisson_pvalue "0.00e+00";
chr4	piRNAome	piRNA_cluster	115292840	115297741	.	+	.	cluster_id "piRO-rno-C00000069"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "23"; poisson_pvalue "0.00e+00";
chr4	piRNAome	piRNA_cluster	123789805	123793246	.	+	.	cluster_id "piRO-rno-C00000070"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "1.02e-09";
chr4	piRNAome	piRNA_cluster	145670017	145678731	.	+	.	cluster_id "piRO-rno-C00000071"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "2.36e-07";
chr4	piRNAome	piRNA_cluster	161151443	161153265	.	+	.	cluster_id "piRO-rno-C00000072"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "17"; poisson_pvalue "0.00e+00";
chr5	piRNAome	piRNA_cluster	91131019	91137363	.	+	.	cluster_id "piRO-rno-C00000073"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "6"; poisson_pvalue "6.31e-09";
chr6	piRNAome	piRNA_cluster	125360395	125363325	.	-	.	cluster_id "piRO-rno-C00000074"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "5"; poisson_pvalue "3.65e-09";
chr7	piRNAome	piRNA_cluster	20125984	20141364	.	-	.	cluster_id "piRO-rno-C00000075"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "17"; poisson_pvalue "0.00e+00";
chr7	piRNAome	piRNA_cluster	20695351	20703370	.	-	.	cluster_id "piRO-rno-C00000076"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "7"; poisson_pvalue "3.82e-09";
chr7	piRNAome	piRNA_cluster	31883002	31883756	.	-	.	cluster_id "piRO-rno-C00000077"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "8"; poisson_pvalue "0.00e+00";
chr7	piRNAome	piRNA_cluster	35626375	35634637	.	-	.	cluster_id "piRO-rno-C00000078"; cluster_type "de_novo"; tier "TIER_1"; n_piRNA "30"; poisson_pvalue "0.00e+00";
chr7	piRNAome	piRNA_cluster	35724063	35729193	.	-	.	cluster_id "piRO-rno-C00000079"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "14"; poisson_pvalue "0.00e+00";
chr7	piRNAome	piRNA_cluster	125460986	125466075	.	-	.	cluster_id "piRO-rno-C00000080"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "8.63e-09";
chr7	piRNAome	piRNA_cluster	144812252	144813057	.	-	.	cluster_id "piRO-rno-C00000081"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "5"; poisson_pvalue "4.12e-11";
chr7	piRNAome	piRNA_cluster	20258730	20263599	.	+	.	cluster_id "piRO-rno-C00000082"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "5"; poisson_pvalue "3.03e-07";
chr7	piRNAome	piRNA_cluster	20319044	20321643	.	+	.	cluster_id "piRO-rno-C00000083"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "1.62e-10";
chr9	piRNAome	piRNA_cluster	114127533	114134950	.	-	.	cluster_id "piRO-rno-C00000084"; cluster_type "de_novo"; tier "TIER_2"; n_piRNA "6"; poisson_pvalue "1.10e-09";
chr9	piRNAome	piRNA_cluster	100728628	100731954	.	+	.	cluster_id "piRO-rno-C00000085"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "5"; poisson_pvalue "1.29e-09";
chrM	piRNAome	piRNA_cluster	1069	7738	.	+	.	cluster_id "piRO-rno-C00000086"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "12"; poisson_pvalue "0.00e+00";
chrX	piRNAome	piRNA_cluster	107250132	107253151	.	-	.	cluster_id "piRO-rno-C00000087"; cluster_type "de_novo"; tier "TIER_3"; n_piRNA "5"; poisson_pvalue "1.89e-15";